Flowering plants are heterosporous, producing two types of spores. Microspores are produced by meiosis inside anthers while megaspores are produced inside ovules, inside an ovary. In fact, anthers typically consist of four microsporangia and an ovule is an integumented megasporangium. Both types of spores develop into gametophytes inside sporangia. As with all heterosporous plants, the gametophytes also develop inside the spores (are endosporic).
A flower is a modified stem tip with compressed internodes, bearing structures that are highly modified leaves.[1] In essence, a flower develops on a modified shoot or axis from a determinate apical meristem (determinate meaning the axis grows to a set size).
Flowers may be directly attached to the plant at their base (sessile--the supporting stalk or stem is highly reduced or absent). The stem or stalk subtending a flower is called a peduncle. If a peduncle supports more than one flower, the stems connecting each flower to the main axis are called pedicels. The apex of a flowering stem forms a terminal swelling which is called the torus or receptacle. The parts of a flower are arranged in whorls on the receptacle. The four main whorls (starting from the base of the flower or lowest node and working upwards) are as follows:
An example of a "perfect flower", this Crateva religiosa flower has both stamens (outer ring) and a pistil (center).
* Calyx: the outermost whorl consisting of units calledsepals; these are typically green and enclose the rest of the flower in the bud stage, however, they can be absent or prominent and petal-like in some species.
* Corolla: the next whorl toward the apex, composed of units called petals, which are typically thin, soft and colored to attract animals that help the process of pollination.
* Androecium (from Greek andros oikia: man's house): the next whorl (sometimes multiplied into several whorls), consisting of units called stamens. Stamens consist of two parts: a stalk called a filament, topped by an anther where pollen is produced by meiosis and eventually dispersed.
* Gynoecium (from Greek gynaikos oikia: woman's house): the innermost whorl of a flower, consisting of one or more units called carpels. The carpel or multiple fused carpels form a hollow structure called an ovary, which produces ovules internally. Ovules are megasporangia and they in turn produce megaspores by meiosis which develop into female gametophytes. These give rise to egg cells. The gynoecium of a flower is also described using an alternative terminology wherein the structure one sees in the innermost whorl (consisting of an ovary, style and stigma) is called a pistil. A pistil may consist of a single carpel or a number of carpels fused together. The sticky tip of the pistil, the stigma, is the receptor of pollen. The supportive stalk, the style, becomes the pathway for pollen tubes to grow from pollen grains adhering to the stigma.
Although the arrangement described above is considered "typical", plant species show a wide variation in floral structure. These modifications have significance in the evolution of flowering plants and are used extensively by botanists to establish relationships among plant species.
In the majority of species, individual flowers have both functional carpels and stamens. These flowers are described by botanists as being perfect or bisexual. Some flowers lack one or the other reproductive organ and called imperfect or unisexual If unisex flowers are found on the same individual plant but in different locations, the species is said to be monoecious. If each type of unisex flower is found only on separate individuals, the plant is dioecious.
Additional discussions on floral modifications from the basic plan are presented in the articles on each of the basic parts of the flower. In those species that have more than one flower on an axis, the collection of flowers is termed an inflorescence. Some inflorescences are composed of many small flowers arranged in a formation that resembles a single flower. The common example of this is most members of the very large composite (Asteraceae) group. A single daisy or sunflower, for example, is not a flower but a flower head—an inflorescence composed of numerous flowers (or florets).
Many flowers have a symmetry, if the perianth is bisected through the central axis from any point, symmetrical halves are produced—the flower is called regular or actinomorphic, e.g. rose or trillium. When flowers are bisected and produce only one line that produces symmetrical halves the flower is said to be irregular or zygomorphic. e.g. snapdragon or most orchids.
Christmas Lillium (Lilium longiflorum). 1. Stigma, 2. Style, 3. Stamens, 4. Filament, 5. Petal
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